Coulson and the agents are thrust backwards in time and stranded in 1931 New York City; with an all-new Zephyr set to time-jump at any moment, the team must hurry to find out exactly what happened.

With the identity of the timeline-unraveling "thread" revealed, the team's mission to protect him at all costs leads each agent to question their own values.

A surprise leap forward in time strands Enoch in 1931 and lands the team in another unfamiliar decade; in order to stop the chronicoms from launching their future-dismantling plan, the agents must infiltrate one of S.H.I.E.L.D.'s most secure bases.

Agent Phil Coulson comes to a realization about the importance of July 22, 1955, in the S.H.I.E.L.D. history books.

The team - Daniel Sousa in tow - reunites with more than one familiar face at the S.H.I.E.L.D. hangout and discovers exactly how to dismantle the Chronicoms' latest plan; the Zephyr unexpectedly leaps forward again.

After blowing its cover and damaging the Zephyr in the process, the team must scramble to rescue Mack's parents, save S.H.I.E.L.D. from a chronicom infestation and fix the ship; Daisy and Sousa are at a disadvantage against Nathaniel Malick.

After getting stranded in 1982, Mack retreats to his childhood home to deal with the death of his parents; Deke scouts a group of new agents.

With the Zephyr's time drive malfunctioning, the team hurtle toward disaster with Yo-Yo as its only hope; Yo-Yo must enlist the help of an old adversary to regain her powers.

A time storm ravages the Zephyr, propelling it toward destruction while simultaneously forcing Daisy and Coulson to relive their failed attempts to save the team.

The team arrives back in 1983, where Nathaniel and Kora are hard at work building an army; Daisy is tasked with protecting Jiaying; the others learn they'll need to face another supercharged enemy to stop Malick.

With help from Kora on the inside, Sibyl and Nathanial continue their fight to shape a dark new future for S.H.I.E.L.D., managing to stay one step ahead of the agents along the way.

Sibyl attempts to get the location of Fitz from Jemma's memories. Meanwhile, the SHIELD team attempts to stop Nathaniel's plans for anarchy and the Chronicoms' attempts to rewrite history.

The final fight against the Chronicoms begins. A familiar face returns.